Eric Thompson has always lived in the shadow of his older brother, the Alpha of the Canadian werewolf pack. While he has strength and speed and a howl that can truly make his pack whole, he doesn't have his brother's viciousness – and the pack wasn't worth either of them dying. To Eric, a leader should lead wisely and well, and his brother has done neither. So he left to become a lone wolf.

Holly Sanchez has likewise lived in the shadows. A full human born to the Omega of the Boulder pack, she was attacked and turned into the very thing she had come to hate. Worse still, she became an alphic healer – except that as a wolf, she now terrifies the very animals she'd hoped to one day treat as a veterinarian.

But as the world goes mad and species begins to attack species, Holly and Eric learn that they may only have their special gifts, and each other, to rely on to hold together the fragments of what was once a mighty shapeshifter population.

For She has finally come, and the world will never again be the same.